List of Pinguicula species

This article is a list of the Butterworts, the species of the genus Pinguicula. For general information on butterworts, please see the main article on Pinguicula.

The genus Pinguicula contains the 80 species of butterworts, belonging to the Bladderwort family (Lentibulariaceae). It has a natural distribution across most of the northern hemisphere, though over half of the species are concentrated in Mexico and Central America. Siegfried Jost Casper systematically divided them into three subgenera with 15 sections. Subsequent phylogenetic research showed that many of these groupings are polyphyletic, but they are used below.

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Subgenus Isoloba

Section Agnata

• Pinguicula agnata
• Pinguicula albida
• Pinguicula filifolia
• Pinguicula gigantea
• Pinguicula benedicta
• Pinguicula cubensis
• Pinguicula infundibuliformis
• Pinguicula pilosa

Section Cardiophyllum

• Pinguicula crystallina
  • Pinguicula crystallina ssp. hirtiflora

Section Discoradix

• Pinguicula casabitoana
• Pinguicula lignicola

Section Heterophyllum

• Pinguicula acuminata
• Pinguicula heterophylla
• Pinguicula kondoi
• Pinguicula mirandae
• Pinguicula parvifolia
• Pinguicula rotundiflora
• Pinguicula imitatrix
• Pinguicula conzattii

Section Isoloba

• Pinguicula lilacina
• Pinguicula pumila: Small Butterwort
• Pinguicula sharpii
• Pinguicula takakii
• Pinguicula caerulea: Blueflower Butterwort
• Pinguicula ionantha: Godfrey's Butterwort, Violet Butterwort
• Pinguicula lutea: Yellow Butterwort
• Pinguicula planifolia: Chapman's Butterwort
• Pinguicula primuliflora: Southern Butterwort
• Pinguicula lusitanica

Subgenus Pinguicula

Section Crassifolia

• Pinguicula esseriana
• Pinguicula ehlersiae
• Pinguicula debbertiana
• Pinguicula jaumavensis

Section Homophyllum

• Pinguicula greenwoodii
• Pinguicula jackii

Section Longitubus

• Pinguicula calderoniae
• Pinguicula crassifolia
• Pinguicula hemiepiphytica
• Pinguicula laueana
• Pinguicula utricularioides

Section Nana

• Pinguicula villosa: Hairy Butterwort

Section Orcheosanthus

• Pinguicula macrophylla
• Pinguicula oblongiloba
• Pinguicula stolonifera
• Pinguicula colimensis
• Pinguicula cyclosecta
• Pinguicula mesophytica
• Pinguicula moranensis
• Pinguicula orchidioides (Synonym P. stolonifera)
• Pinguicula potosiensis
• Pinguicula rectifolia
• Pinguicula zecheri
• Pinguicula gypsicola
• Pinguicula moctezumae
• Pinguicula elizabethiae

Section Orchidioides

• Pinguicula laxifolia

Section Pinguicula

• Pinguicula balcanica
  • Pinguicula balcanica var. tenuilaciniata
• Pinguicula corsica

• Pinguicula poldinii
• Pinguicula grandiflora
  • Pinguicula grandiflora ssp. rosea
• Pinguicula leptoceras
• Pinguicula longifolia
  • Pinguicula longifolia ssp. causensis
  • Pinguicula longifolia ssp. dertosensis
  • Pinguicula longifolia ssp. reichenbachiana
• Pinguicula macroceras: California Butterwort
  • Pinguicula macroceras var. macroceras
  • Pinguicula macroceras ssp. nortensis
• Pinguicula mundi
• Pinguicula nevadensis
• Pinguicula vallisneriifolia
• Pinguicula vulgaris: Common Butterwort

Subgenus Temnoceras

Section Ampullipalatum

• Pinguicula antarctica
• Pinguicula calyptrata
• Pinguicula chilensis
• Pinguicula involuta
• Pinguicula elongata

Section Micranthus

• Pinguicula algida
• Pinguicula alpina : Alpine Butterwort

• Pinguicula ramosa
• Pinguicula variegata

Section Temnoceras

• Pinguicula clivorum
• Pinguicula crenatiloba
• Pinguicula emarginata
• Pinguicula gracilis
• Pinguicula immaculata

References, External links, and Further Reading

See the main Pinguicula article

• Casper, S. J.: Monographie der Gattung Pinguicula, (Bibliotheca Botanica, Heft 127/128), 1966, Stuttgart
• Cieslak, Thomas; Polepalli, Jai Santosh; White, Adam; Müller, Kai; Borsch, Thomas; Barthlott, Wilhelm; Steiger, Juerg; Marchant, Adam; Legendre, Laurent: Phylogenetic analysis of 'Pinguicula (Lentibulariaceae): chloroplast DNA sequences and morphology support several geographically distinct radiations, 2005, Am. J. Bot. 92: 1723 - 1736 Abstract
• Schlauer, Jan: Carnivorous Plant Database, version Nov. 15, 16:25.